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Greenhouse ventilation

I've a small heated greenhouse with a few veg. and bonsai plants taken inside for the winter. Due to the weather it's been shut up quite tightly over the past week or so. I always keep the greenhouse well ventilated in summer but just wondered about winter. When its not freezing should I open a window or would the plants be happier being kept a bit warmer with everything shut up.
I recall reading an article about the importance of greenhouse ventilation but can't find it now.
At about 750 feet on the western edge of The Pennines. Clay soil.
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When you don't even know who's in the team
S.Yorkshire/Derbyshire border
I open my greenhouse door every day; the vents are always left slightly open. Of course on really inaccesible days I may leave it.
You can do your own test by visiting your greenhouse after about 1 hour of full sun. A max-min thermometer is essential, summer and winter.
"Have nothing in your garden that you don't know to be useful, or believe to be beautiful."